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Couplet 143
143a: Hind tarsus with 5 tarsomeres, basal tarsomere reduced, often difficult to see, either hidden in apical excavation of hind tibia or shorter than fourth length of second tarsomere and obliquely attached beneath it (may only be visible from below in oblique distal angle, see Figure 26-10D); elytra covering pygidium Bostrichidae p.426
143b: First tarsomere usually not so reduced; if so, then pronotum not hood-like, head not hypognathous or hind tarsus of 4 tarsomeres, pygidium exposed
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