Couplet 23
23a: Elytra shortened and widely divergent at apex (except in Lichnanthe lupina), not covering pygidium; eighth morphological abdominal segment with spiracle Glaphyridae p. 411
23b: Elytra not shortened or widely divergent at apex, pygidium exposed or not (Figures 26-29C,D, 26-30, 26-32, 26-33, 26-34); eighth morphological abdominal segment lacking spiracle Scarabaeidae p. 412
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